Misc Pap 191

MISCELLANEOUS PAPERS

Centre for Entomological Studies Ankara

•k'k'k'k'k'k'k'k'k'k'k'k'k'k'k'k'k'k'k'k'k'k'k'k'k'k'k'k'k'k'k'k'k'k'k'k'k 'k'k'k'k’k'k'k'k'k'k'k'k'k'k'k'k'k'kJc'k'k'k'k'k'k'k'k'k'k'k'k'k'k'k'k'k

No 191 26 May 2019 ISSN 1015-8235

http://zoobank.org/References/EA25F218-A260-4623-A98E-32B6C7E4A7BB

On the small Heterocera collection from Palawan Island
(Philippines), with some remarks

Muhabbet Kemal 1 Sibel Kizildag Ahmet Omer Ko^ak

Abstract: On the small Heterocera collection from Palawan Island (Philippines), with some
remarks. Misc. Pap. 191: 1-26, 52 figs.

This paper deals with the Heterocera material from Olangu-an Mt. (North Palawan, Philippines).
Totally 88 species of 17 families are listed and faunistically evaluated. Remarks on various subjects
are added to some species. Generally, the information provided is supported by original images.
Key words: Heterocera, Lepidoptera, Aganaidae, Arctiidae, Cossidae, Ctenuchidae, Ethmiidae,
Geometridae, Hyblaeidae, Lasiocampidae, Lymantriidae, Limacodidae, Noctuidae, Notodontidae,
Oecophoridae, Pyralidae, Sphingidae, Thyridae, Uraniidae,fauna, Palawan, Philippines.

Among the Lepidoptera of the Philippines stored in the Cesa Collection, a small number of
the Heterocera from Mt. Olangu-an (North Palawan Island, Philippines) are prepared here for a
faunistic paper (Fig. 1 ).

The authors have been carrying out studies on the Heterocera fauna of the Philippines from
various standpoints (Kogak & Kemal, 2011; Kemal, Kizildag & Kogak, 2018, 2019a; Kemal & Kogak,
2019).

First comprehensive publication on the Heterocera of Palawan Island belongs to
Pagenstecher (1890a). He listed 93 taxa from this island 2 . Semper’s (1896-1902) monumental work
on the Lepidoptera (Heterocera) of the Philippines cannot be ignored here.

In the present paper 88 species of 17 families are listed from Olangu-an Mt. (Palawan,
Philippines), together with their original references and ranges as codes (Kogak & Kemal, 2019). A
few rare species from Mt. Gantung and Mt. Mantalingajan from Palawan are also evaluated here.
Remarks to some species are added. At the end of the list, two comments on the latest paper of the
authors on the Lepidoptera of the Caragan Mt. are given.

All the material studied are identified by the authors, by using especially the Cesa Reference
Collection. For this purpose, the following publications are also worth to note: Cerny (2011),
Hogenes & Treadaway (1998), Holloway (1983-2005), Kononenko & Pinratana (2005, 2013),
Schintlmeister & Lourens (2010), Seitz (ed.) (1907-1934), Zolotuhin et al. (1998). The material and
the related documents are currently preserved in the Cesa Collection and Archive 3 .

1 Dr Muhabbet Kemal & Dr Sibel Kizildag - Van Yiiziincii Yil University, Faculty of Science, Dept, of Biology, Campus, Van / Turkey,
e-mails: muhabbet kemal( 5 )yahoo.com.tr & sibelkizildag( 5 )yyu.edu.tr

Prof, em Dr Ahmet Omer Kogak, c/o Yiiziincii Yil University, Faculty of Science, Dept, of Biology., Van / Turkey,
e-mail: cesa tr( 5 )yahoo.com.tr

2 Currently 468 Heterocera species are known from Palawan Island (Cesa Info-system: http://www.cesa-tr.org/Infos.htm )

3 http: / / entcesa.tripod.com/Cesacollection.pdf

ISSN 1015-8235

Miscellaneous Papers
Centre for Entomological Studies Ankara

Nr 191 -26 05 2019

Fig. l - Overview of some Heterocera specimens in envelope from Mt. Olangu-an (North Palawan,
Philippines) in the Collection of the Cesa.

List of the species

The Heterocera species from Mt. Olangu-an [Olg] are listed below alphabetically. The symbol
* denotes that the mtCOI DNA of the species will be examined separately.

Aganaidae

i Agape chloropyga (Walker, 1854 )

Original reference: H ypsa chloropyga Walker,1854, List Spec, lepid. Insects Colin Br. Mus. 2: 454-
455 - Type: Australia.

Range tin code!: A U CN Yu MY MM TH CMa Int Knc VN ID Bor Snd Mlc Aru Bor Flo Ja Sum Crm CIb PG NBr PH
Luz Mno Crg Pal Olg

Material studied : i<$. Philippines, North Palawan, Mt. Olangu-an, January, 1998, F. Mamigo leg.
(Cesa)

Remarks : This species is widely distributed species but the present record seems first for the fauna
of Palawan Island.

zAsota egens (Walker, 1854 ) (Fig. 6 )

Original reference: Hypsa egens Walker,1854, List Specimens lepid. Insects Colin Br. Mus. 2: 453.

Range (in code): IN Adm NP TH Knc CMa Dst MY Bor SG TW ID Bli Ja Sum Egg Nia Clb Tim Snd Wt JP CN TW PG
PH Pal Olg Mno

Material studied : 5$ (wingspan 5i-6omm). Philippines, North Palawan, Mt. Olangu-an, January,
1998, F. Mamigo leg. (Cesa)

3 Asota heliconia (Linnaeus, 1758 )

Original reference: P halaena heliconia Linnaeus,1758, Syst. Nat. (Edn.10) 1: 511. "Habitat in
Calidis regionibus".

Range fin code): MM Tns TH CMa Int Knc CN TW JP Ri ID Egg Tim PH Pal Olg Bal Ley Blc Mno Crg Neg Boh Luz
Pq Sam Sib AU PG Bor Sbh KBa Nia Snd Bli Aru Amb Crm Adm Bsm Nk Ja Clb

Material studied : 2c? (typical and f. nervosa ). Philippines, North Palawan, Mt. Olangu-an, January,
1998, F. Mamigo leg. (Cesa)

2

Heterocera collection from northern Palawan ...

ISSN 1015-8235

Miscellaneous Papers
Centre for Entomological Studies Ankara

Nr 191 -26 05 2019

4A.sotajavana (Cramer,[1780]) (Fig. 7)

Original reference: P halaena javana Cramer,[1780], Uitl. Kapellen 3: 146, 175, pl.274 fig.C. Type:
Java.

Range fin code!: T H Cbi Ntt Hba MM MY Bor KBa Swk Sbh ID Ja Clb Flo Tim KH PH Ley BIc Luz Ifg Mdo Mno Apo
Crg Busa Sam Pal Olg

Material studied : 1$. Philippines, North Palawan, Mt. Olangu-an, January, 1998, F. Mamigo leg.
(Cesa)

sA.sota heliconioides (Moore,1878) (Fig. 8)

Original reference: N eochera heliconioides Moore,1878, Proc. zool. Soc. Lond. 1878: 6. Type S'
Philippines, Luzon (coll. Stgr.) (BMNH).

Range (in code!: ID Ja PH Bal Luz Mno Dav Mdo Pal Olg

Material studied : 2c?. Philippines, North Palawan, Mt. Olangu-an, January, 1998, F. Mamigo leg.
(Cesa)

6 .Euplocia membliaria (Cramer,[1780]) (Fig. 9)

Original reference: P halaena membliaria Cramer,[1780], Uitl. Kapellen 3:139,175.

Range (in code): IN Adm MM TH Knc ID MY Bor Sum Clb Snd Ja Flo Nat PH Pan Bly Luz Pq Pal Olg

Material studied : 1 S- Philippines, North Palawan, Mt. Olangu-an, January, 1998, F. Mamigo leg.
(Cesa)

Arctiidae

7 Amerila astreus (Drury,1773)

Original reference: S phinx astreus Drury,1773, Illust. Nat. Hist. Exot. Insects 2: index, 49 pl.28
fig.4. Type: Bengal.

Range (in code): PK IN Ben Mh Nsk Bor MY ID Sum Ja PH Luz Pq Mar Sam Ceb Boh Cmt Mno Neg Mpl Pal Olg PG
BD CN Hn SZ Yu Kwa TW MM

Material studied : lS- Philippines, North Palawan, Mt. Olangu-an, January, 1998, F. Mamigo leg.
(Cesa)

8 .Pareuchaetes pseudoinsulata Rego Barros,i956

Original reference: P areuchaetes pseudoinsulata Rego Barros,i956, Revta Bras. Ent. 6: 79. Type:
Venezuela, Caracas.

Range (in code): V E Crc TT LKTH MY Bor GU PH Luz Mar Mdo Pan Ley Mno Pal Olg Boh

Material studied : 4c?. Philippines, North Palawan, Mt. Olangu-an, January, 1998, F. Mamigo leg.
(Cesa)

Remarks : This species is reported here from Bohol Island for the first time (Cerny, 2011).

9 .Lemyra maculifascia (Walker,1855)

Original reference: S pilosoma maculifascia Walker,1855, List Specimens lepid. Insects Colin Br.
Mus. 3: 676. Type: Java.

Range (in code): ID Ja Bli TH CMa Cuo Cu04 PG AU PH Luz Ifg Mar Pan Sam Ley Mno Pal Olg Boh

Material studied : 5c?. Philippines, North Palawan, Mt. Olangu-an, January, 1998, F. Mamigo leg.
(Cesa)

10 .Spilosoma strigatula Walker,1855

Original reference: S pilosoma strigatula Walker,1855, List Specimens lepid. Insects Colin Br. Mus.
3: 613. Type: Burma.

Range (in code): CN MM TH ID PH Luz Mdo Pal Olg Boh Mno Crg

Material studied : lc?. Philippines, North Palawan, Mt. Olangu-an, January, 1998, F. Mamigo leg.
(Cesa)

3

Heterocera collection from northern Palawan ...

ISSN 1015-8235

Miscellaneous Papers
Centre for Entomological Studies Ankara

Nr 191 -26 05 2019

Cossidae

11 *Zeuzera cqffeae Nietner,l86i

Original reference :Zeuzera coffeae Nietner,i86i, Enemies of Coffee Tree: 21-22.

Range fin code! : PK IN Up Kma Rnb LKTH Bnk Nan Phy MY Bor TW ID Mlc Krkt PG Ij Ja PH Ley Blc Pal Olg

Material studied : lS- Philippines, North Palawan, Mt. Olangu-an, January, 1998, F. Mamigo leg.
(Cesa).

Remarks : This species is apparently new for the fauna of Palawan. A male specimen from Leyte
Island (Balacaue Mt.) is also found in the Cesa Collection.

Ctenuchidae

12.*? Amata sp. (Fig. 10)

Material: 2c?. Philippines, North Palawan, Mt. Olangu-an, January, 1998, F. Mamigo leg. (Cesa)
Remarks : Identification process of this specimen is ongoing.

Ethmiidae

*Ethmia palaivana Schultze,i925 (Fig. 11)

Original reference: E thmia palaivana Schultze,i925, Philippine J. Sci. 28: 574. Type $\ Palawan
(lost) (Diakonoff, [1968]).

Range (in code) : TH Yal MY Bor BN Sbh PH Pal Mjn Taw

Material studied : 1 $. Philippines, Palawan, Mt. Mantalingajan 1000m [Mjn], 3-103.1998, F.
Mamigo leg. (Cesa)

Geometridae

13 AAchrosis sp.

Material : 9c? Philippines, North Palawan, Mt. Olangu-an, January, 1998, F. Mamigo leg. (Cesa)
Remarks : Identification process of this specimen is ongoing.

14 .*Agathia asterias Meyrick,i888 (Fig. 12)

Original reference: A gathia asterias Meyrick,i888, Proc. Linn. Soc. N.S.Wales 2 (2): 899. Syntypes
c?$: Australia: Queensland, Northern Territories (BMNH).

Range (in code): a u qau nau ph Mdo Pal olg

Material studied : id 1 . Philippines, North Palawan, Mt. Olangu-an, January, 1998, F. Mamigo leg.
(Cesa)

Remarks : The name asterias Meyrick is considered by Holloway 4 5 as a distinct species, or synonym
of Agathia pisina by various authors. The Mindoro record of this species (coll. Sommerer, ZSM) s
was illustrated in the Bold Systems. The present record of this species is new for the fauna of
Palawan.

lS. 'Amblychia sp. (Fig. 13)

Material studied : 1$ (wingspan 60mm), 1$ (wingspan 70mm). Philippines, North Palawan, Mt.
Olangu-an, January, 1998, F. Mamigo leg. (Cesa)

Remarks : Identification process of the specimens is ongoing.

4 Holloway, J.D., 1977. Lepidoptera of Norfolk Island. Their Biogeography and Ecology.

5 http://v51.boldsvstems.org/index.php/Public SearchTerms?querv=%22Agathia%2oasterias%22rtaxl

4

Heterocera collection from northern Palawan ...

ISSN 1015-8235

Miscellaneous Papers
Centre for Entomological Studies Ankara

Nr 191 -26 05 2019

16. Biston subflavus Inoue,i985 (Fig. 14)

Original reference: B iston subflavus Inoue,i985, Cho Cho 8 (11): 22, 24, figs. 3, 5, 7. Holotype f:
Philippines, Negros Is., Amlam.

Range (in code!: PH Neg Ami Knl Ley Blc Pan Bly Pal Olg

Material studied : 6 ff. Philippines, North Palawan, Mt. Olangu-an, January, 1998, F. Mamigo leg.
(Cesa)

Remarks : This species is new to the fauna of Palawan Island.

17 . Bracca georgiata (Guenee,[i858]> (Fig. 15)

Original reference: P anaethia georgiata Guenee,[i858], Hist. nat. Insectes, Spec. Gen. Lep. 10:
196. Holotype $: "Borneo" (coll. Gn.) [Genitalia of the Holotype was illustrated by Holloway
(fig- 357 )]-

Range (in code): MY Bor ID Crm Nat Ja Sgr PH Mdo Luz Pq Mno Pal Olg TH Ntt Hba

Material studied : if. Philippines, North Palawan, Mt. Olangu-an, January, 1998, F. Mamigo leg.
(Cesa)

Remarks : This species is reported and illustrated by the authors from Caragan Mt (Mindanao)
(Kemal, et al., 2019b).

18. *Celerenapalawanica Pagenstecher,i890 (Fig. 16)

Original reference : Celerena palawanica Pagenstecher,i890, Dt. ent. Z., Iris 3: 31. Syntypes if 1$:
[Philippines]: Palawan.

Range (in code) : ph Pal oig BaiMdo

Material studied : 6 f 4$ (wingspan 52-560101). Philippines, North Palawan, Mt. Olangu-an,
January, 1998, F. Mamigo leg. (Cesa)

Remarks : There is also another species, Celerena spodoptera, described by Inoue (1999). Our
specimens belong to palawanica Pagenstecher externally. This species has not been examined so
far from the molecular standpoint.

*Craspedosis stigmatias Herbulot,1985 (Fig. 17)

Original reference: Craspedosis stigmatias Herbulot,i985, Nouv. Revue Ent. (N.S.) 2 (3): 258, figs.
10,17. Holotype f: Palawan (coll. Herbulot).

Material studied : if (wingspan 42mm). Philippines, South Palawan, Mt. Gantung 1000m, F.
Mamigo leg. (Cesa)

Remarks : This ennomid species was described by Herbulot from Central Palawan, Mt. Salokot and
endemic for this island. It is illustrated here because of its importance.

19 *?Chrysocraspeda sp. (Fig. 18)

Material : if (wingspan 21mm). Philippines, North Palawan, Mt. Olangu-an, January, 1998, F.
Mamigo leg. (Cesa)

Remarks : The identity of the genus is temporary. The process is still ongoing.

20.Dyspha.nia malayanus (Guerin-Meneville,i843) (Figs 2,19)

Original reference: Hazis malayanus Guerin-Meneville,i843, Delessertes Souv. Vy. l'lnde 2: 89.

Range (in Code): MY IN SI Djl Bor Swk Prk PH Pal Olg ID Sum

Material studied : 1$ (GP3113) (wingspan 60mm). Philippines, North Palawan, Mt. Olangu-an,
January, 1998, F. Mamigo leg. (Cesa)

Remarks : There is no significant genetic distance (0.30-0.42%) between the populations Palawan
and Sumatra/Myanmar from Boldsystems (Ratnasingham & Hebert, 2007).

21 *Dysphania plena (Walker,1856) (Fig. 20)

Original reference: E uschema plena Walker,1856, List Specimens Lepid. Ins. Colin Br. Mus. 7:
1668. Type: Philippines.

Range (in code): ph Luz Mnl Pal olg

Material studied : 6c? (wingspan 66-78mm). Philippines, North Palawan, Mt. Olangu-an, January,
1998, F. Mamigo leg. (Cesa)

Remarks : Indo-Malayan Dysphania species are well illustrated by Yazaki (1989).

5

Heterocera collection from northern Palawan...

ISSN 1015-8235 Miscellaneous Papers Nr 191 -26 05 2019

Centre for Entomological Studies Ankara

Fig. 2 - Dysphania malayanus (Geometridae). Female genitalia (GP3113). Philippines, Palawan, Mt. Olangu-an.

22*?Hyperythra sp. (Fig. 21)

Material : Philippines, North Palawan, Mt. Olangu-an, January, 1998, F. Mamigo leg. (Cesa)

Remarks : The identity of the genus is temporary. The process is still ongoing.

23. Hyposid.ra talaca (Walker,i860)

Original reference: Lagyra talaca Walker,i860, List Specimens Lepid. Insects Colin Br. Mus. 20:
59. Syntypes $$: Celebes, Philippine Isles, Ceylon.

Range (in code): ID Crm Clb LK IN PH Pal Olg AU QAu SB TH CMa Cuo Cmz Czio Knc Cbi Phe Ntt Hba MY Bor
Material studied : i( 3 i$. Philippines, North Palawan, Mt. Olangu-an, January, 1998, F. Mamigo leg.
(Cesa)

24 . Pareumelea hortensiata (Guenee,[i858]) (Fig. 22)

Original reference: E umelea hortensiata Guenee,[i858], Hist. nat. Ins., Spec. gen. Lep. 9: 394.
Range (in code): my sg id sum th sip cib ph Mdo Pal oig

Material studied : if. Philippines, North Palawan, Mt. Olangu-an, January, 1998, F. Mamigo leg.
(Cesa)

25 . Problepsis apollinaria (Guenee,[i858])

Original reference: A rayris apollinaria Guenee,[i 8581 , Hist. nat. Insectes, Spec. Gen. Lep. 10: 13,
555 (index). Type $: "Borneo", "coll. Gn."

Range (in code): MY Bor Swk PG Bsm IN ID Kai PH Luz Mno Crg Neg Mpl Pal Olg TH AU

Material studied : 5$. Philippines, North Palawan, Mt. Olangu-an, January, 1998, F. Mamigo leg.
(Cesa)

26 . Pseudeuchromia maculifera (R.Felder & Rogenhofer,i875) (Fig. 23)

Original reference: B ursada maculifera R.Felder & Rogenhofer,i875, Reise Fregatte Novara, Bd 2
(Abth.2) (5): pi. 130 fig.12. Type: Moluccas.

Range (in code): ID Mlc Clb PH Neg Mpl Knl Mno Luz Pan Bly Pal Olg MY Bor Sbh KBa

Material studied : 46'$. Philippines, North Palawan, Mt. Olangu-an, January, 1998, F. Mamigo leg.
(Cesa)

27. Ruttelleronapallicostaria (Moore,[1868])

Original reference: A ngerona pallicostaria Moore,[1868], Proc. zool. Soc. Lond. 1867: 620. Type:
"Bengal".

Range (in code): IN Ben BT TH Ntt Hba CMa Cuo Cuoi Fn Nna Kao Cbi Knc Nst PH Neg Pal Olg MY Bor ID Sum

Material studied : 5$. Philippines, North Palawan, Mt. Olangu-an, January, 1998, F. Mamigo leg.
(Cesa)

28. *?Serratophyga sp.

Material studied : 1 ex. Philippines, North Palawan, Mt. Olangu-an, January, 1998, F. Mamigo leg.
(Cesa)

Remarks : The identity of the genus is temporary. The process is still ongoing.

6

Heterocera collection from northern Palawan ...

ISSN 1015-8235

Miscellaneous Papers
Centre for Entomological Studies Ankara

Nr 191 -26 05 2019

Hyblaeidae

29.Hyblaea constellata Guenee,i852 (Fig. 24)

Range (in code): IN Mh Nsk LK Dck Pdy Kndy CN JP TW MM TH AU QAu PG ID Sum ?PH ?Pal ? 01 g

Material studied : 4 f $. Philippines, North Palawan, Mt. Olangu-an, January, 1998, F. Mamigo leg.
(Cesa)

Remarks : Apparently new to the fauna of the Philippines.

Lasiocampidae

30 .Lebeda intermedia Holloway,1987 (Fig. 25)

Original reference: L ebeda intermedia Holloway,1987, Moths of Borneo 3: 21-22, pl.9, fig.10.
Holotype A Borneo, Kuching (BMNH).

Range (in COdek PH Pal Olg Mno Luz ID Sum MY Bor

Material studied : if. Philippines, North Palawan, Mt. Olangu-an, January, 1998, F. Mamigo leg.
(Cesa)

Remarks : This species was decribed by Holloway from Borneo. Zolotuhin et al. (1998) reported this
species from the Philippines (Palawan).

*Odonestis erectilinea ssp .frater Sergeev & Zolotuhin,2010 (Fig. 26)

Original reference : Odonestis erectilinea ssp. frater Sergeev & Zolotuhin,2010, Atalanta 41 (3/4):
389, figs. Holotype f: Philippinen: Palawan N., Mt. Cleopatra needle 6oo-75om, 2i.xi- 2.xii. 2000,
leg. D.Mohagan (MWM).

Range (in code) : phP alGnt

Material studied : if. Philippines, South Palawan, Mt. Gantung 1000m [Gnt], 10-28.ii.1998, F.
Mamigo leg. (Cesa).

Our older specimen of this endemic taxon is here illustrated.

31 .Suana concolor (Walker,1855)

Original reference : Lebeda concolor Walker, 1855, List Specimens lepid. Insects Colin Br. Mus. 6:

1463.

Range (in code) : IN BD Syl LK MM CN VN Ton MY PH Luz Ifg Mdo Hlc Pan Neg Knl Ley Blc Sam Boh Mno Bkd
Ktd Mtu Apo Bal Pal Olg Ceb Taw ID Sum Bor Ja Bnk TH CMa Cmz Czio Fn Int Nan Ntt Hba Kbi

Material studied : if. Philippines, North Palawan, Mt. Olangu-an, January, 1998, F. Mamigo leg.
(Cesa)

Remarks : This species was reported by Hogenes & Treadaway (1998: 53) as “Palawan, Ulanguan”.

Lymantriidae
32.Aroa sp.

Material studied : if. Philippines, North Palawan, Mt. Olangu-an, January, 1998, F. Mamigo leg.
(Cesa)

33 *Carriola sp.

Material studied : if. Philippines, North Palawan, Mt. Olangu-an, January, 1998, F. Mamigo leg.
(Cesa)

Remarks : The members of this species are sexually dimorph. There are several similar species in
the region; therefore, its identity is ongoing.

34 *Cozola austriaca (Semper,1899) (Fig. 27)

Original reference: E uproctis austriaca Semper,1899, Reisen Archipel. Philipp. 2: 469, pl.54 fig.6.
Syntypes 2$: Philippines, Mindanao.

Range (in code): ph Mno Pal oig

Material studied : 1$. Philippines, North Palawan, Mt. Olangu-an, January, 1998, F. Mamigo leg.
(Cesa)

7

Heterocera collection from northern Palawan...

ISSN 1015-8235

Miscellaneous Papers
Centre for Entomological Studies Ankara

Nr 191 -26 05 2019

35 .Olene mendosa Hubner,[i823]

Original reference: Olene mendosa Hiibner,[i823], Samml. exot. Schmett. Zutr. 2: 19, nr. 147, figs.
293-294.

Range fin code!: LK Kndy Dck Pdy Cob IN BD Syl ID Ja CN TW AU PG VN Ton Tda TH Bnk CMa Cuo Cuos CDo
CD2 CRa Loe Knc Cbi MY Bor PH Mno Crg Pal Olg

Material studied : lc?. Philippines, North Palawan, Mt. Olangu-an, January, 1998, F. Mamigo leg.
(Cesa)

Limacodidae

36 .*Cheromettia sp. (Fig. 28)

Material studied : lc?. Philippines, North Palawan, Mt. Olangu-an, January, 1998, F. Mamigo leg.
(Cesa)

Remarks : Apparently, it looks like a Cheromettia species, but identification process is still ongoing.

Noctuidae

37. A nomis sp.

Material studied : 2 ex. Philippines, North Palawan, Mt. Olangu-an, January, 1998, F. Mamigo leg.
(Cesa)

Remarks : Identity process is still ongoing.

38 . *Bocula marginata (Moore,1882)

Original reference: B orsippa marginata Moore,1882, [in] Hewitson & Moore, Descr. Indian lepid.
Insects Colin late Atkinson: 173. Type: "Darjiling".

Range (in code) : IN SI Djl TH Knc CMa Cmz Czii Int LA NP CN TW PH Pal Olg

Material studied : lc?. Philippines, North Palawan, Mt. Olangu-an, January, 1998, F. Mamigo leg.
(Cesa)

Remarks : This specimen looks like very much to this species externally (Kononenko & Pinratana,
2005); however, it carries no median transversal lines on the upperside of the forewing. Apparently
new for the fauna of the Philippines.

39 . Calesia stillifera R.Felder & Rogenhofer,i874

Original reference: Calesia stillifera R.Felder & Rogenhofer,i874, Reise Fregatte Novara, Bd 2
(Abth.2) (5): pi. cxvii fig. 18.

Range (in Code): T H CMa Int Cmz Czll Sbu KH VN IN LK CN PH Luz Mnl Pal Olg

Material studied : i<$. Philippines, North Palawan, Mt. Olangu-an, January, 1998, F. Mamigo leg.
(Cesa)

Remarks : This species is new for the fauna of Palawan Island.

40 . Callyna monoleuca Walker,1858

Range (in code): Z A IN Up Bhi CN TW TH Knc CMa Lmp MY PH Mno Pal Olg ID Clb JP AU

Material studied : 2 ( 5 '. Philippines, North Palawan, Mt. Olangu-an, January, 1998, F. Mamigo leg.
(Cesa)

Remarks : New to the fauna of Palawan Island.

41 .Chrysopera combinans (Walker,1858) (Fig. 29)

Range (in Code): LK ID Ja Clb TH CMa Int my in np cn fj SB PH Pal Olg AU

Material studied : 4c? $. Philippines, North Palawan, Mt. Olangu-an, January, 1998, F. Mamigo leg.
(Cesa)

Remarks : Pagenstecher (1890b: 109-110) described “Achaea quadrilunata” from East Java
(Indonesia), which is currently treated as a junior synonym of Achaea combinans Walker,1858 in
the genus Chrysopera Hampson,i894. This species has not been reported from Palawan so far
(Pagenstecher, 1890a).

New record for the fauna of Palawan Island.

8

Heterocera collection from northern Palawan...

ISSN 1015-8235

Miscellaneous Papers
Centre for Entomological Studies Ankara

Nr 191 -26 05 2019

42. *Episteme vetula (Geyer,i832) (Fig. 30)

Original reference: H eraclia vetula Geyer,i832, Zutr. Samml. exot. Schmett. 4: 17-18, figs. 657-
658. Type: Java.

Range fin code!: ID Nia Sum Ja Bor Swk Tim BD Syl Sbh KBa Sdk PH Pal Olg TH CRa CMa Dst MM MY Pen Bor
KHVN

Material studied : lS (wingspan 52mm). Philippines, North Palawan, Mt. Olangu-an, January,
1998, F. Mamigo leg. (Cesa)

Remarks : This record belongs to palavardca Jordan,1912 ([in] Seitz, A. Gross-Schmett. Erde 11: 3),
originally described at subspecific level. The genetic distance between Palawan and Malaysia
populations will be prepared soon. Another species from Palawan is *Exsula victrix
(Westwood,1848), represented in the Cesa Collection by a single male from Mt. Gantung 1000m
(Palawan) (Fig. 31) (New record for the fauna of the Philippines). This specimen shows some
discrepancies when we compare it by using the descriptions given by Jordan (op. cit., 9).

43 . Eudocima homaena (Hiibner,[i823])

Original reference: Othreis homaena Hiibner,[i823], Verz. bekannt. Schmett.: 264.

Range (in code): T H Phr Wgc CMa Int MY VN NP LK IN Mh Nsk ID Bor Tim Flo PH ?Pal ? 01 g CN JP TW

Material studied : 2 ex. Philippines, North Palawan, Mt. Olangu-an, January, 1998, F. Mamigo leg.
(Cesa)

44 . Hamodes propitia (Boisduval,i832) (Fig. 32)

Original reference \ Ophiusa propitia Boisduval,i832, [in] d'Urville, Voy. Astrolabe 1: 244. Type:
Nouvelle-Irlande.

Range (in Code) :PG NIr Bsm TH CMa Int Dst MY LA MM IN Mh Nsk LK NP BD CN TW ID PH Pal Olg

Material studied : l$. Philippines, North Palawan, Mt. Olangu-an, January, 1998, F. Mamigo leg.
(Cesa)

Remarks : The present record seems first for the fauna of Palawan Island.

45 . Hypopyra ossigera Guenee,i852 (Fig. 33)

Range (in code) : TH Pgg CMa Int MY VN CN Yu IN SI Ass Ben TW ID Sum Clb PH Pal Olg

Material studied : lS (wingspan 80mm). Philippines, North Palawan, Mt. Olangu-an, January,

1998, F. Mamigo leg. (Cesa)

Remarks : This record is new for the fauna of the Philippines (Kononenko & Pinratana, 2005).

46 .Ischyja infema Swinhoe,i902 (Fig. 34)

Original reference: I schyja infema Swinhoe,i902, Ann. Mag. nat. Hist. (7) 10: 498. Type S'- "Kina
Balu, N. Borneo (Waterstradt).

Range (in code): PH Pal Olg Bor Sbh KBa MY TH Tng CMa Int ID Sum Ja Lb Clb IN

Material studied : lS- Philippines, North Palawan, Mt. Olangu-an, January, 1998, F. Mamigo leg.
(Cesa)

47 . Masca abactalis Walker, [1859]

Original reference: M asca abactalis Walker,[1859], List Specimens lepid. Insects Colin Br. Mus. 16
(1858): 9.

Range (in code): SG MY Prk Bor Sbh TH Knc CN Yu ID Sum Ja Amb Tim Flo Clb Mlc Crm Wg Ij PG PH Mno Crg Pal
Olg JP MM VN

Material studied : 6c? $. Philippines, North Palawan, Mt. Olangu-an, January, 1998, F. Mamigo leg.
(Cesa)

48 . Mecodina praecipua (Walker,1865) (Fig. 35)

Original reference: Thermesia praecipua Walker,1865, List Specimens lepid. Insects Colin Br.
Mus. 33:1056. Type $: "Ceylon".

Range (in code): IN N 1 Adm LK ID Amb Ja Tim Ij TH Cbi Nrt Kao PG PH Pal Olg AU JP

Material studied : lS- Philippines, North Palawan, Mt. Olangu-an, January, 1998, F. Mamigo leg.
(Cesa)

Remarks : This record is new for the fauna Palawan.

9

Heterocera collection from northern Palawan ...

ISSN 1015-8235

Miscellaneous Papers
Centre for Entomological Studies Ankara

Nr 191 -26 05 2019

49 .Mods undata (Fabricius,i775)

Original reference \Noctua undata Fabricius,i775, Syst. Ent.: 6oo. Type(s): India orientalis.

Range tin code!: PKK-P SLCM ZRKE ZWHar MURUSb Us CNTW JP INMhNskLKMM AdmNkID Ja Bor Swk
Sda Sum Tim Flo Sbw PG PC ZA Nta Kzu Dbn TH CMa Cuo Cuoi Knc Rng KHVNNPPH Boh Mno Crg Pal Olg

Material studied : 2c?. Philippines, North Palawan, Mt. Olangu-an, January, 1998, F. Mamigo leg.
(Cesa)

Remarks : The present record seems first for the fauna of Palawan Island.

50 .Ommatophora luminosa (Cramer,1780) (Fig. 36)

Original reference : Phalaena luminosa Cramer,1780, Uitl. Kapellen 3:147, pl.274 fig.D. Type:
[Indonesia]: Java.

Range fin code]: TH Tng CMa Int CN IN Ass MM KH VN TW LK MY Bor PH Mno Bkd Ktd Pal Olg ID Ja Sum Ja Bli
Flo Clb PG

Material studied : 1 ex. Philippines, North Palawan, Mt. Olangu-an, January, 1998, F. Mamigo leg.
(Cesa)

Remarks : The present record seems first for the fauna of Palawan Island.

51 .Ophthalmis lincea (Cramer,[1779])

Original reference: P halaena lincea Cramer,[1779], Uitl. Kapellen 3: 61, pl.228 Fig.B. Type:
Amboina.

Range (in code): ID Amb Crm Bia SB PH Bal Neg Knl Pal Olg MY Bor PG NIr

Material studied : 4c?. Philippines, North Palawan, Mt. Olangu-an, January, 1998, F. Mamigo leg.
(Cesa)

sz.Oxyodes scrobiculata (Fabricius,i775)

Original reference: N octua scrobiculata Fabricius,i775, Syst. Ent.: 252. Type(s): "India Orientalii".

Range (in code): T H CMa Cuo Cuoi CUO7 Czio Cmz CDo Dst DS09 Int Phr Nrt Knc Sbu Bnk Cph Cbi Pht Cbu MY
Bor VN LK Dck Pdy IN NP CN TW JP KP KR ID Sum Sba Tim Flo PG WS NH NC F J PH Mno Crg Bkd Ktd Pal Olg

Material studied : 8c?. Philippines, North Palawan, Mt. Olangu-an, January, 1998, F. Mamigo leg.
(Cesa)

53 .Platyja umminia (Cramer,[1780]) (Fig. 37)

Original reference : Phalaena umminia Cramer,[1780], Uitl. Kapellen 3 (22): 117, (23-24): pl.267
fig.F. Type: Java, Samarang.

Range (in code): ID Clb Ja MY CN HK TH Knc Nrt Kao VN LA MM IN BD LK Hn TW Bor Sbw Adm Mlc PG PH Pal
Olg Luz Pq Pan Bly AU QAu GU

Material studied : 3c?. Philippines, North Palawan, Mt. Olangu-an, January, 1998, F. Mamigo leg.
(Cesa)

54 .Spirama helicina (Hiibner,i824) (Fig. 38)

Original reference: S peiredonia helicina Hiibner,i824, Zutr. Samml. exot. Schmett. 3: pl.76 fig.437.

Range (in code): JP TH Nrt Kao CMa Int Cuo CUO7 VN IN NP ID Ja Sum PH Neg Knl Pal Olg CN TW KP KR RU Us

Material studied : i<$. Philippines, North Palawan, Mt. Olangu-an, January, 1998, F. Mamigo leg.
(Cesa)

55 .Spirama retorta (Clerk,1759) (Fig. 39)

Range (in code): T H CMa Cmz Czio Czll Int CDo Php MM VN KH NP IN PK BD LK Pdy ID CN TW JP KP KR PH
Luz Mnl Pal Olg Ch BD Syl

Material studied : 5 A Philippines, North Palawan, Mt. Olangu-an, January, 1998, F. Mamigo leg.
(Cesa)

56 .Sympis rufibasis Guenee,i852 (Fig. 40)

Original reference: S ympis rufibasis Guenee,i852, Hist. nat. Ins., Spec. gen. Lepid. 7 (Noct 3): 344,
pl.24 fig.i. Type: "Java".

Range (in code): TH Knc Kph CMa CDo CD2 Cmz Czio Pap Mro Nrt Sbu Nna Che Cbi Phc Trat Cph Tng LA KH VN
MM NP IN LK CN TW PH Ley Blc Pal Olg ID Ja Bor Sum Bli Tim Flo Sbw Clb Amb Mlc Crm SB AU QAu PG

Material studied : lc?. Philippines, North Palawan, Mt. Olangu-an, January, 1998, F. Mamigo leg.
(Cesa)

10

Heterocera collection from northern Palawan ...

ISSN 1015-8235

Miscellaneous Papers
Centre for Entomological Studies Ankara

Nr 191 -26 05 2019

57 . Tirumala sp.

Material studied : i<$. Philippines, North Palawan, Mt. Olangu-an, January, 1998, F. Mamigo leg.
(Cesa)

58 . Tiruvaca subcostalis (Walker,1865) (Fig. 41)

Range fin code!: PG MY IN TH Tng CMa Int MM ID Sum Ja Bor CIb SB PH Boh Pal Olg

Material studied : id 1 . Philippines, North Palawan, Mt. Olangu-an, January, 1998, F. Mamigo leg.
(Cesa)

Remarks : New record for the fauna of Palawan Island.

59 . Westermannia triangularis Moore,1877 (Fig.42)

Original reference : Westermannia triangularis Moore, 1877, Proc. zool. Soc. London 1877 (3): 605-
606. Type: "S.Andamans" (BMNH).

Range (in code): IN Adm MY Bor Sbh ID Kit Sum Clb PH Pal Olg

Material studied : id 1 . Philippines, North Palawan, Mt. Olangu-an, January, 1998, F. Mamigo leg.
(Cesa)

Remarks : This species is new for the fauna of Palawan Island.

Notodontidae

6o.Gargetta hampsoni Schintlmeister,i98i (Fig. 43)

Original reference : Gargetta hampsoni Schintlmeister,i98i, Atalanta 12: 286-289, figs. Holotype
< 3 : Indonesia, Sumatra.

Range (in code): ID Sum TH CMa CDo MY Bor PH Mno Dav Apo Busa Neg Knl Mdo Hlc Pal Olg

Material studied : 5$. Philippines, North Palawan, Mt. Olangu-an, January, 1998, F. Mamigo leg.
(Cesa)

61 .Phalera sundana Holloway,1982 (Figs 3, 44)

Original reference : Phalera sundana Holloway,1982, Appendix, [in] Barlow,H., An introduction to
the moths of South East Asia: 201, pl.21 fig.5. Holotype S\ Malaysia, W.Pahang, Genting Tea Estate
(BMNH).

Range (in code) : ID Sum Ja Bli CN MY PH Ley Sam Ceb Neg Mar Luz Mno Pal Olg

Material studied : 1$ (GP3114) (wingspan 79mm). Philippines, North Palawan, Mt. Olangu-an,
January, 1998, F. Mamigo leg. (Cesa)

Remarks : This species described from Malaysia but distributed also some islands of Indonesia and
the Philippines (Schintlmeister & Lourens,20io). The male genitalia is also very variable (shapes of
uncus and accessory spines on the valva) in various islands of the Philippines. Shapes of socii
remarkably variable. It is interesting that the tetraxon spines in the aedeagus are numerous in
some notodontids ( Semidonta, Mesophalera, Fusadonta, etc), including Phalera sundana. They
may be transferred to the bursa copulatrix of the female during copulation 6 .

62 .Rodneya cernyi Schintlmeister,1993 (Fig. 45)

Original reference: Rodney a cernyi Schintlmeister,1993, Nachr. ent. Ver., Apollo, Suppl. 12: 133-
134, figs. Holotype S'- Philippinen, S-Palawan, Mt. Gantung.

Range (in code): PH Luz Pal Gnt Olg Mar Mdo Neg Mno LA TH

Material studied : 6A Philippines, North Palawan, Mt. Olangu-an, January, 1998, F. Mamigo leg.
(Cesa)

6 cf. Chapman,T.A., 1913. On the shedding of the cornuti in Pyrausta aurata, Sc. Entomologist's Rec. J. Var. 25:106-108, 4 pis.

11

Heterocera collection from northern Palawan...

ISSN 1015-8235

Miscellaneous Papers
Centre for Entomological Studies Ankara

Nr 191 -26 05 2019

Fig. 3 — Phalera sundana (Notodontidae). Male genitalia (GP3114) with separately enlarged spines. Philippines,
Palawan, Mt. Olangu-an.

63 .Spatalia (Celeia) sikkima (Moore,1879) (Figs 4, 5)

Original reference : Celeia sikkima Moore,1879, Descr. Indian lep. Atkinson (1): 63. Type S'-
"Darjiling".

Range fin code! : IN SI Djl NP Adm MM ID Ja Clb Sum MY CN Hun VN Ann Ton Bao M-c PH Pal Olg Luz Mdo Pan
Sam Ley Mno

Material studied : 5S (GP3116). Philippines, North Palawan, Mt. Olangu-an, January, 1998, F.
Mamigo leg. (Cesa)

Remarks : Particularly the shape of the aedeagus, uncus and the socii of the male genitalia indicates
that it belongs to Spatalia sikkima Moore (cf. Schintlmeister & Lourens,20io). This species was
reported from the Philippines (incl. Palawan) by Schintlmeister and Lourens (2010). Abdominal
processes are illustrated here for the first time.

Oecophoridae
*Tonica sp.

Range fin code) : PHPaiMjn

Material studied : 1$. Philippines, Palawan, Mt. Mantalingajan 1000m [Mjn], 3-103.1998, F.
Mamigo leg. (Cesa)

Remarks : This genus is new to the fauna of Palawan (Diakonoff, [1968]).

12

Heterocera collection from northern Palawan ...

ISSN 1015-8235

Miscellaneous Papers
Centre for Entomological Studies Ankara

Nr 191 -26 05 2019

Figs 4, 5 - Spatalia sikkima (Notodontidae). Male genitalia and abdominal segments with lateral processes (GP3116).
Philippines, Palawan, Mt. Olangu-an.

Pyralidae

64. Endocrossisflavibasa.lis (Moore,1867)

Original reference: B otyodes flavibasalis Moore,1867, Proc. zool. Soc. London 1867: 96. Syntypes
India "Bengal".

Range tin code!: PH Luz Mno Crg Pal Olg IN Ass Ben SI BT NP CN Kwa TH Tng MM Prk MY PG PMo ID Sum Nia

Material studied : i< 3 . Philippines, North Palawan, Mt. Olangu-an, January, 1998, F. Mamigo leg.
(Cesa)

65. Eth.opia gigantea Owada,i986 (Fig. 46)

Original reference: E thopia gigantea Owada,i986, Kontyu 54 (4): 618-623, 8 figs. Holotype S'-
Philippines, Palawan, 1985, ex Y.Nishiyama (NSMT).

Range tin code!: ph PalMjn oig

Material studied : lS (wingspan 72mm). Philippines, North Palawan, Mt. Olangu-an, January,
1998, F. Mamigo leg. (Cesa)

Remarks : This species was previously reported by the authors from southern Palawan Island (Mt.
Mantalingajan) (Kogak & Kemal, 2011).

66. Haritalodes derogata (Fabricius,i775)

Original reference: Phalaena derogata Fabricius,i775, Syst. Ent.: 641. Type(s): India orientalis.

Range (in code): ZA AU TH CMa Cuo CUO7 CDo CD2 Sbu Bnk Sak NP PK MM BD SG AF Srb IR Gh LK MY Bor VN
ID Ja CN KP TW JP PH Pal Olg RU Sb WS FJ NC SB PG IN OW SC RE KM MG ZR GM GH MW NG SL UG TZ ZM ZW

Material studied : 1 $. Philippines, North Palawan, Mt. Olangu-an, January, 1998, F. Mamigo leg.
(Cesa)

67 . Nevrina procopia (Stoll, [1781])

Original reference: P halaena procopia Stoll,[1781], [in] Cramer, Uitl. Kapellen 4 (29-31): 152,
pl.368 fig.E.

Range (in code): A C JP TW PH Luz Pq Ley Pao CmM Mno Crg Pal Olg IN BT LK Nia Ja Sgr Amb ID PG CN MY SG

Material studied : 1 $. Philippines, North Palawan, Mt. Olangu-an, January, 1998, F. Mamigo leg.
(Cesa)

13

Heterocera collection from northern Palawan...

ISSN 1015-8235

Miscellaneous Papers
Centre for Entomological Studies Ankara

Nr 191 -26 05 2019

68 .Omphisa anastomosalis (Guenee,i854)

Original reference : Pionea ? anastomosalis Guenee,i854, [in] Boisduval & Guenee, Hist. nat. Ins.,
Spec. gen. Lepid. 8: 373-374. Type: "Java".

Range tin code]: MM TH Bnk CN LK IN Adm ID Ja D-YTW Hw VNKHLA JP PH Luz Pal Olg PG D-Y

Material studied : i<$. Philippines, North Palawan, Mt. Olangu-an, January, 1998, F. Mamigo leg.
(Cesa)

69. Pygospila tyres (Cramer,[1780])

Original reference: P halaena tyres Cramer,[1780], Uitl. Kapellen 3: pl.263 fig.C.

Range fin code]: PK Cp PH Boh Luz Pq CmM Neg Mpl Mno Pal Olg IN NP LK MM TH CMa Cuo Cu07 Cmz Czio Bor
Ja Grm Clb Arab Aru PG AU ID ZA CM ZR Cl MW NG SL ZW

Material studied : 4 ( 5 '. Philippines, North Palawan, Mt. Olangu-an, January, 1998, F. Mamigo leg.
(Cesa)

7 0. Vitessa (pyraliata-gr.) splendida Schultze,i9o8 (Fig.47)

Original reference: Vitessa splendida Schultze,i9o8, Philipp. J. Sci. 3(1) A: 35, pl.i f.11. Holotype $
[destroyed]: Philippines: Negros occ.: Maao.

Range (in Code): PH Luz Mnl Mno Neg ?Pal ?oig

Material studied : 1 ex. Philippines, North Palawan, Mt. Olangu-an, January, 1998, F. Mamigo leg.
(Cesa)

71 .Vitessa {stettina- gr.) philippina Munroe & Shaffer,1980 (Fig.48)

Original reference : Vitessa philippina Munroe & Shaffer,1980, Bull. Br. Mus. nat. Hist. (Ent.) 39
(4): 286-287, figs- Holotype $: Philippines: Luzon: Manila (Wileman) (BMNH).

Range (in Code) : PH Luz Mnl Mno Dav Apo ?Pal ?oig

Material studied : 1 ex. Philippines, North Palawan, Mt. Olangu-an, January, 1998, F. Mamigo leg.
(Cesa)

Remarks : The genus Vitessa is represented by two specimens in our collection, belonging to two
distinct species. Their temporary identities are based upon the external morphology (Munroe &
Shaffer,1980). In their revisional work, the new species, philippina was based upon the female
only. The species splendida was described by Schultze in 1908, basing upon the female from
Negros Island (now destroyed). It is currently known from Luzon, Negros, Mindanao, and a single
female from Palawan in the Zoological Museum, Copenhagen. Male descriptions of both species
have not been published so far. Therefore, it is hard to decide about their precise identities. More
material or DNA studies seems to be ineffective on the solution.

Sphingidae

72A.cosm.eryx anceus (Stoll,[1781])

Original reference: S phinx anceus Stoll,[1781], [in] Cramer, Uitl. Kapellen 4 (29-31): 124, pl.ccclv.
Type "Amboina".

Range (in code): LK Kndy IN Hp Meg Ass Khs NP MM TH CMa Cuo CDo MY PG AU VN ID Sum Ja Sba Clb Flo Amb
PH Bal Boh Cal Ceb Din Jol Ley Luz Mar Mdo Mno Neg Pal Olg Pan Sam Sga Stu Taw

Material studied : 3V (wingspan 62-70mm). Philippines, North Palawan, Mt. Olangu-an, January,
1998, F. Mamigo leg. (Cesa)

73 Acosmeryx pseudonaga Butler,1881

Original reference: A cosmeryx pseudonag a Butler,1881, Ill. typical Spec. Lep. Het. Colin Br. Mus.
5: 2, pi. lxxviii fig.3. Type: "Bhotan". Range (in code): bt th my cn vn id Sum Ja Bor ph Pal olg Bal
Material studied : iV (wingspan 82-84mm). Philippines, North Palawan, Mt. Olangu-an, January,
1998, F. Mamigo leg. (Cesa)

74Ambulyx moorei Moore, [1858]

Original reference: A mbulyx moorei Moore,[1858], [in] Horsfield & Moore, Cat. lep. Ins. Mus. East
India Coy 1: 266 [proposed in synonymy and without description]. Range (in code): ph Bal Pal olg in

LK Adm MY TH CMa Cuo Bor Sum Ja VN CN Hn

14

Heterocera collection from northern Palawan ...

ISSN 1015-8235

Miscellaneous Papers
Centre for Entomological Studies Ankara

Nr 191 -26 05 2019

Material studied : 3c? (wingspan 86-92mm). Philippines, North Palawan, Mt. Olangu-an, January,
1998, F. Mamigo leg. (Cesa)

75 . Cechenena helops (Walker,1856) (Fig. 49)

Original reference: P hilampelus helops Walker, 1856, List Specimens Lepid. Ins. Colin Br. Mus. 8:
180. Type: Port Natal [incorrect] "Malaysia, Penang" (Pittaway & Kitching]. Range (in code!: np in

TH VN MY Pen ID Sum Ja Bor Clb Mlc Crm PG AU PH Bal Din Ley Luz Mar Mdo Hlc Mno Apo Dav Neg Pal Olg Pan
Sam Siq

Material studied : 3c? (wingspan 90-98mm). Philippines, North Palawan, Mt. Olangu-an, January,
1998, F. Mamigo leg. (Cesa)

76 . Eupanacra busiris (Walker,1856)

Original reference: P anacra busiris Walker,1856, List Specimens Lepid. Ins. Colin Br. Mus. 8: 158-

159 - Type(s): "Silhet". Range (in code): IN BD Syl MM NP THCNVNMY ID Sum Ja Bor Clb Adm PH Bal Pal
Olg Ley Blc Mdo Mno

Remarks : This species was reported by Hogenes & Treadaway (1998: 57) as “Palawan, Ulanguan”.

T7.Hippotion celerio (Linnaeus,1758)

Original reference: S phinx celerio Linnaeus,1758, Syst. Nat. (Edn.10) 1: 491.

Range (in code): M d cn ma dz tn ly pt es fr Co sa si it mt ie gb be nl lu de ch at pl cz sk hu ro yu al

BG Slv Vrn GRRdCrTRCYLB Bei IL Hf Pa IR Kh Th Gls Ss DK RU MD UA BY GG Smt Bjo IN Tn LK ID Sum Ja Kai
AU QAu Hw PG NC NH FJ ZA Fs Cap Bio Tva Lpo Spb Mdk MZ GN SA SL PH Bal Boh Cal Ceb Dum Jol Ley Luz Pq Mar
Mdo Mno Neg Pal Olg Pan Pao Sam Tic WS TO EL LO SB MY Gau Pre Mpu Brb Jh Nta Kzu Dbn SL JO AF Sfk Srb Nu
Bsg PK Swa Sgl TH CMa Cuo TW Cc Cp Abe Rbt Btna Agi Orn Blid Mzg Hrh 31 33 35 48

Material studied : lc? (wingspan 64mm). Philippines, North Palawan, Mt. Olangu-an, January,
1998, F. Mamigo leg. (Cesa)

78. Macroglossumfa.ro (Cramer,1780)

Original reference: S phinx faro Cramer,1780, Uitl. Kapellen 3:165, pl.285 fig.2.

Range (in code): CN IN TH VN TW JP Ri MY ID Sum Ja Bor PH Ley Pal Olg

Remarks : This species was reported by Hogenes & Treadaway (1998: 81) as “Palawan, Ulanguan”.

79 . Macroglossumjani Hogenes & Treadaway,1998

Original reference: M acroglossum jani Hogenes & Treadaway,1998, Nachr. Ent. Ver. Apollo
(Suppl.) 17: 90-92. Holotype <$: Philippines, Palawan, Languan, 25.ii.1995 (CCGT) (Paratypes from
Homonhon, Ulanguan, Cebu).

Range (in code): ph Ceb Hom Pal olg

Remarks : This species was reported by Hogenes & Treadaway (1998: 90) as “Palawan, Ulanguan”.

80 . Marumba tigrinus Gehlen,i936 (Fig. 50)

Original reference: M arumba tigrina Gehlen,i936, Ent. Z., Frankf. a. M. 49: 461, fig.

Range (in Code): ID Sum Ja Bor PH Pal Olg TH

Material studied : 5c? (wingspan 88-9imm). Philippines, North Palawan, Mt. Olangu-an, January,
1998, F. Mamigo leg. (Cesa)

8 1. Megacorma obliqua (Walker,1856) (Fig. 51)

Original reference: M acrosila obliqua Walker,1856, List Specimens Lepid. Insects Colin Br. Mus.
8: 208. Type: "Ceylon".

Range (in code): IN LK MM TH CMa Cuo VN MY ID Sum Bor Ja PH Bal Jol Ley Luz Mno Neg Pal Olg Sam Cm PG
SB

Material studied : 3c? (wingspan H2-n8mm) ?. Philippines, North Palawan, Mt. Olangu-an,
January, 1998, F. Mamigo leg. (Cesa)

8z.Meganoton rufescens (Butler,[1876])

Original reference: Diludia rufescens Butler,[1876], Proc. zool. Soc. Lond. 1875: 260.

Range (in code): IN TH CMa CDo CN MM MY Adm VN AU QAu PH Ley Luz Mno Pan Sam Pal Olg ID Cm Amb Hm
Ja Sum Bor PG

7 Perhaps the biggest sphingid species in the Philippines is Amplypterus panopus ssp. mindanaoensis Inoue,i996. Wingspan of 3 c? $
varies from 133 to 151mm (from Negros I., Kanlaon Mt. 1500m, 12-17.xii.1997, F.Mamigo leg. (Cesa)).

15

Heterocera collection from northern Palawan ...

ISSN 1015-8235

Miscellaneous Papers
Centre for Entomological Studies Ankara

Nr 191 -26 05 2019

Material studied : 2A Philippines, North Palawan, Mt. Olangu-an, January, 1998, F. Mamigo leg.
(Cesa)

Ss.Theretra clotho (Drury,[1773])

Original reference: S phinx clotho Drury,[1773], Illust. Nat. Hist. Exot. Insects 2: 48.

Range fin codel: LK IN NP TH CMa Cuo CNKPKRJP Adm ID Sly Mlc AU PG NC Sum Ja Flo Tim TW PH Bal Boh
Bon Cal Ceb Jol Ley Mdo Mno Neg Pal Olg Pan Sam Sib Siq Stu Taw

Material studied : 3 c? (wingspan 68-84mm). Philippines, North Palawan, Mt. Olangu-an, January,
1998, F. Mamigo leg. (Cesa)

S^.Theretra nessus (Drury,[1773])

Original reference: S phinx nessus Drury,[1773], Illust. exot. Ins. 2: Index [91], pl.76, fig.i.

Range (in code): LK Pdy IN NP MM TH CMa Cuo Sgk Hya PSu VN CN JP TW AU QAu NH LO NC Hw PH Bal Boh
Cal Cat Ceb Din Jol Ley Luz Mar Mas Mdo Mno Neg Knl Pal Olg Pan Sam Sib Siq Stu

Material studied : 4c? (wingspan 97-i02mm). Philippines, North Palawan, Mt. Olangu-an, January,
1998, F. Mamigo leg. (Cesa)

Ss.Theretra suffusa (Walker,1856)

Original reference: Chaerocampa suffusa Walker,1856, List Specimens lepid. Insects Colin Br.
Mus. 8:146.

Range (in Code): N P IN MM TH CN VN TW JP Ri MY ID Sum Ja Bor PH Pal Olg

Remarks : This species was reported by Hogenes & Treadaway (1998: 53) as “Palawan, Ulanguan”.

Thyridae

(=Thyrididae auct.)

86 *Canaea sp.

Material studied : 2c?l$. Philippines, North Palawan, Mt. Olangu-an, January, 1998, F. Mamigo
leg. (Cesa)

Remarks : The identity of the specimens is ongoing. The genus has not been reported from the
Philippines so far (Whalley,i97i).

Uraniidae

87 . Decetia pallidaria Pagenstecher,i888 (Fig. 52)

Original reference: Decetia (?) pallidaria Pagenstecher,i888, Jb. nassau. Ver. Naturk. 41:166-167.
Syntypes c?$: Amboina.

Range (in code) : id mic cib my Bor Swk ph Pal oig

Material studied : 2c?. Philippines, North Palawan, Mt. Olangu-an, January, 1998, F. Mamigo leg.
(Cesa)

Remarks : This species was reported from Palawan by Pagenstecher (1890: 26).

88 . Micronia aculeata Guenee,[i858]

Original reference: M icronia aculeata Guenee,[i858], Hist. nat. Insectes, Spec. Gen. Lep. 10: 22-
24. Type-species: Micronia aculeata Guenee,[i858].

Range (in Code): CN TW IN LK MM Adm MY Bor ID Ja Clb PH Pal Olg

Material studied : if. Philippines, North Palawan, Mt. Olangu-an, January, 1998, F. Mamigo leg.
(Cesa)

16

Heterocera collection from northern Palawan ...

ISSN 1015-8235

Miscellaneous Papers
Centre for Entomological Studies Ankara

Nr 191 -26 05 2019

Additional information to the paper : Kemal,M., Kizildag,S. & A.O.Koqak, 2019. On the
Heterocera of Mt. Caragan (Philippines, Mindanao), with some remarks. Misc Pap. 190: 1-
50 .

On page 5, nr.12. Abraxas sp.? n.: This species may be near to Abraxas gephyra West, 1929 (Novit.
Zool. 35: 124), described from the Philippines, Luzon (Baguio) but without illustration. The
identification of this species is still ongoing.

On page 26, nr. 76. Athetis maculatra, described by Lower from Australia (Queensland), was cited
from Caragan Mt. with hesitation. After preparing the barcode (6s8bp) of the specimen from
Caragan Mt. (barcoded by the Cesa (Lep-Noctoog)) and comparing with the Australian A.
maculatra specimen from Bold Systems, the genetic distance was calculated as 0.8 between them.
It means that the specimen from Caragan Mt. is merely a different population of Athetis maculatra
(Lower,1902). Consequently, Athetis maculatra (Lower) is reported from the Philippines for the
first time.

Acknowledgements

We sincerely thank to Mr. Katsumi Yazaki (Tokyo, Japan) for his kind help in supplying some
important publications.

References

Cerny, K., 2011. A Review of the subfamily Arctiinae (Lepidoptera: Arctiidae) from the Philippines.
Entomofauna 32 (3): 29-92.

Diakonoff,A. [1968]. Microlepidoptera of the Philippine Islands. U.S. Nat. Mus. Bull. 257:1-483.
Hogenes,W. & C.G.Treadaway, 1998. The Sphingidae (Lepidoptera) of the Philippines. Nachr. ent. Ver.
Apollo Suppl. 17:17-132, figs.

Holloway, J.D., 1983-2005, The Moths of Borneo, Pts 4-18. Kuala Lumpur.

Inoue,H., 1999. A new species of the genus Celerena Walker (Lepidoptera, Geometridae) from Palawan,
Philippines. Tinea, 16 (1): 1-3.

Kemal,M., Kizildag,S. & A.O.Ko^ak, 2018. A preliminary list of the moth species of Mindanao Island
(Philippines), in the Cesa Collection, with some remarks (Lepidoptera). Cesa News 160: 1-15, 29 figs.
Kemal,M., Kizildag,S. & A.O.Ko^ak, 2019a. Some Heterocera of Bohol Island (Philippines), a
preliminary list of the species in the Cesa collection with faunistical notes. Misc. Pap. 182:1-20, 52 figs.
Kemal,M., Kizildag,S. & A.O.Ko^ak, 2019b. On the Heterocera of Mt. Caragan (Philippines, Mindanao),
with some remarks. Misc Pap. 190:1-50, 76 figs, 5 maps.

Kemal,M. & A.O.Ko^ak, 2019. A Taxonomical notes on two species of the Philippines (Lepidoptera,
Geometridae). Misc. Pap. 185:1-2.

Ko^ak,A.O. & M. Kemal, 2006. Notes on the nomenclature of some species group taxa in the family
Geometridae (Lepidoptera). Misc. Pap. 102-103:15-16.

Ko^ak,A.O. & M.Kemal, 2011. Ethopia gigantea Owada in Palawan Island, The Philippines C Lepidoptera,
Pyralidae ). Cesa News 69:1-2, 2 figs.

Ko^ak,A.O. & M.Kemal, 2019. First updated geographical codes used in the publications of the Cesa.
Priamus 17 (4): 319-360.

Kononenko,V.S. & B.A.Pinratana, 2005. Moths of Thailand, vol.3. Noctuidae (Part I). 261 pp., 45 Pis.
Bangkok.

Kononenko V. S. & A. Pinratana, 2013. Moth of Thailand Vol. 3, Part 2. Noctuoidea. An illustrated
Catalogue of Erebidae, Nolidae, Euteliidae and Noctuidae (Insecta, Lepidoptera) in Thailand. 625 pp.
Brothers of St. Gabriel in Thailand. Bangkok.

Munroe,E.G. & M. Shaeffer, 1980. A revision of Vitessidia Rothschild & Jordan and Vitessa Moore
(Lepidoptera, Pyralidae). Bull. Br. Mus. Nat. Hist. (Ent.) 39 (4): 241-360.

Owada, M., 1986. A new Galleriine moth of Ethopia (Lepidoptera, Pyralidae) from Palawan, the
Philippines. Kontyu Tokyo 54 (4): 618-623.

17

Heterocera collection from northern Palawan...

ISSN 1015-8235

Miscellaneous Papers
Centre for Entomological Studies Ankara

Nr 191 -26 05 2019

Pagenstecher, A., 1888. Beitrage zur Lepidopteren-Fauna des Malayischen Archipels (V). Verzeichniss der
Schmetterlinge von Amboina. Jb. Nassau. Ver. Naturk. 41: 85-217.

Pagenstecher, A., 1890a. Heteroceren der Insel Palawan. Dt. Ent. Z. Iris 3: 1-33.

Pagenstecher, A., 1890b. Beitrage zur Lepidopteren-Fauna des Malayischen Archipels (VI). Uber
Schmetterlinge von Ost-Java. Jb. Nassau. Ver. Naturk. 43: 93-110.

Ratnasingham,S. & P.D.N.Hebert, 2007. The Barcode of Life Data System ( www.barcodinglife.org ).
Molecular Ecology Notes 7: 355-364.

Robinson,G.S., Tuck,K.R. & M.Shaffer, 1994. A field guide to the smaller moths of south-east Asia.

306 pp. 51 figs., 31 pis. Malaysian Nature Society, Kuala Lumpur.

Schintlmeister, A. & J. Lourens, 2010. The Philippine Notodontidae (Lepidoptera). Quadrifirta 8: 1-349,
figs.

Seitz, A. (ed.), 1907-1934. Die Indo-Australischen Spinner und Schwarmer, Bd.10 (Text & Tafeln), [in] Die
Gross-Schmetterlinge der Erde. Alfred Kernen Verlag, Stuttgart.

Semper,G., 1896-1902. Die Schmetterlinge der philippinischen Inseln. Beitrag zur Indo-Malayischen
Lepidopterenfauna, [in] Semper,C., Reisen im Archipel der Philippinen 6 (2): Die Nachtfalter,
Heterocera , Pis. C.W.Kreidel’s Vlg. Wiesbaden.

Sergeev, A.I. & V. V. Zolotuhin, 2010. A review of the genus Odonestis Germar, 1812 with descriptions of
two new species and one new subspecies (Lepidoptera, Lasiocampidae). Atalanta 41 (3/4): 375-396, figs.
Whalley, P.E.S., 1971. A revision of the genus Canaea Walker (Lepidoptera, Thyrididae). Bull. Br. Mus.
Nat. Hist. (Ent.) 26:161-180, Pis.

Yazaki,K., 1989. An illustrated guide to Dysphania (Lepidoptera, Geometridae). Gekkan-Mushi 217: 2-8,
24 figs.

Zolotuhin,V.V., Treadaway,C.G. &T.J.Witt, 1998. The Lasiocampidae (Lepidoptera) of the
Philippines. Nachr. ent. Ver. Apollo , Suppl. 17:133-222, figs.

18

Heterocera collection from northern Palawan...

ISSN 1015-8235

Miscellaneous Papers
Centre for Entomological Studies Ankara

Nr 191 -26 05 2019

Figures

All the specimens illustrated below are from Mt. Olangu-an (Palawan Island)

Figs 8, 9 -Asota heliconioides (Aganaidae) (left). Euplocia membliaria (Aganaidae) (right).

Figs 10, li — ? Amata sp. (Ctenuchidae) (enlarged) (left). Ethmia palawana (Ethmiidae) (right).

19

Heterocera collection from northern Palawan ...

ISSN 1015-8235

Miscellaneous Papers
Centre for Entomological Studies Ankara

Nr 191 -26 05 2019

Figs 12,13 —Agathia asterias (Geometridae) (left). Amblychia sp.$ (Geometridae, Ennominae) (reduced size) (right).

Figs 14,15 - Bistort subflavus (Geometridae, Ennominae) (left). Bracca georgiata (Geometridae, Ennominae) (right).

Figs 16,17 - Celererta palawanica (Geometridae) (left). Craspedosis stigmatias (Geometridae) (right).

20

Heterocera collection from northern Palawan ...

ISSN 1015-8235

Miscellaneous Papers
Centre for Entomological Studies Ankara

Nr 191 -26 05 2019

Figs 18,19 -? Chrysocraspeda sp. (Geometridae) (left). Dysphania malayanus (Geometridae) (right), for the female
genitalia, see fig. 2.

Figs 22, 23 -Pareumelea hortensiata (Geometridae) (left). Pseudeuchromia maculifera (Geometridae) (right).

21

Heterocera collection from northern Palawan ...

ISSN 1015-8235

Miscellaneous Papers
Centre for Entomological Studies Ankara

Nr 191 -26 05 2019

Figs 24, 25 - Hyblaea constellata (Hyblaeidae) (left). Lebeda intermedia (Lasiocampidae) (right).

Figs 26, 27 —Odonestis erectilinea ssp .frater (Lasiocampidae) (left). Cozola austriaca (Lymantriidae) (right).

Figs 28, 29 - Cheromettia sp. (Limacodidae) (left). Chrysopera combinans (Noctuidae) (right).

22

Heterocera collection from northern Palawan ...

ISSN 1015-8235

Miscellaneous Papers
Centre for Entomological Studies Ankara

Nr 191 -26 05 2019

Figs 30, 31 - Episteme vetula ssp. palavanica (Noctuidae, Agaristinae) (left). Exsula victrix (Noctuidae, Agaristinae)
(right).

Figs 32,33 - Hamodes propitia (Noctuidae) (left). Hypopyra ossigera (Noctuidae) (right).

Figs 34,35 - Ischyja infema (Noctuidae) (left). Mecodina praepicua (Noctuidae) (right).

23

Heterocera collection from northern Palawan ...

ISSN 1015-8235

Miscellaneous Papers
Centre for Entomological Studies Ankara

Nr 191 -26 05 2019

Figs 36, 37 - Ommatophora luminosa (Noctuidae) (left). Platyja umminia (Noctuidae) (right).

Figs 38, 39 - Spirama helicina (Noctuidae) (left). Spirama retorta (Noctuidae) (right).

Figs 40, 41 - Sympis rufibasis (Noctuidae) (left). Tiruvaca subcostalis (Noctuidae) (right).

24

Heterocera collection from northern Palawan ...

ISSN 1015-8235

Miscellaneous Papers Nr 191 - 26 05 2019

Centre for Entomological Studies Ankara

Figs 42, 43 - Westermannia triangularis (Noctuidae) (left). Gargetta hampsoni (Notodontidae) (right).

Figs 44, 45 - Phalera sundana (Notodontidae) (left). Rodney a cernyi (Notodontidae) (right).

Fig. 46 - Ethopia gigantea (Pyralidae, Galleriinae).

Figs 47, 48 - Vitessa splendida (Pyralidae) (left). Vitessa philippina (Pyralidae) (right).

25

Heterocera collection from northern Palawan ...

ISSN 1015-8235

Miscellaneous Papers
Centre for Entomological Studies Ankara

Nr 191 -26 05 2019

Figs 49, 50 - Cechenena helops (Sphingidae) (left). Marumba tigrinus (Sphingidae) (right).

Figs 51, 52 - Megacorma obliqua (Sphingidae) (left). Decetia pallidaria (Uraniidae, Auzeinae) (right).

26

Heterocera collection from northern Palawan ...

ISSN 1015-8235

Miscellaneous Papers
Centre for Entomological Studies Ankara

Nr 191 -26 05 2019

MISCELLANEOUS PAPERS ISSN 1015-8235

Miscellaneous Papers is a peer reviewed serial of the Centre for Entomological Studies Ankara, established in 1989. It appears at
irregular interval in a year, as PDF format and includes original articles of the research workers of the Centre, regarding on various
subjects on Entomology (taxonomy, nomenclature, checklist, fauna, biodiversity, distribution, biogeography, ecology, insect-plant
interactions, bionomy, and behaviour).

The CESA is a non-profit group, no royalties will be paid to authors of contributions. Papers accepted become the copyright of the
related serial.

Miscellaneous Papers is currently archived online at “Internet Archive”, in accordance with the publication rules of the ICZN. It is an
open-access serial, distributed under the terms of the “Creative Commons Attribution License”, which permits free use, and distribution
in any medium, provided the original author(s) and source are credited.

centre -for CiAstoiAAsDlDPjtcPil studies, AAtzava

(A scientific Consortium)

(co-operation of research workers for pure-scientific, not commercial purpose)

Web Page of the Cesa: http://www.cesa-tr.org/

Scientific Serials; Priamus & Priamus Supplement (print and online versions) (ISSN 1015-8243) 8 , Miscellaneous Papers (print and
online versions) (ISSN 1015-8235) 9 , Memoirs (print and online versions) (ISSN-8227) 10 DVD Films 11 , Iconographia Insectorum 12
(online), Cesa Publications on African Lepidoptera (online) 1 ^, Cesa News (online)^, Cesa Books (online) ^

Owners / Sahipleri - Editors / Yayincilar: Prof, em Dr Ahmet Omer Kogak (c/o Van Yiiziincii Yil University, Van, Turkey), Asst.
Prof. Dr Muhabbet Kemal Kogak (c/o Van Yiiziincii Yil University, Van, Turkey).

Editorial Board of all Scientific Serials of the CESA / Biitiin Bilimsel Yayinlann Yayin Kurulu: Insecta, taxonomy,
nomenclature, ecology, faunistics: Prof, em Dr Ahmet Omer Kogak (c/o Van Yiiziincii Yil Universitesi, Turkey), Asst. Prof. Dr Muhabbet
Kemal Kogak (Van Yiiziincii Yil University, Turkey).

Chief referees of all Scientific Serials of the CESA: Prof, em Dr Ahmet Omer Kogak & Asst. Prof. Dr Muhabbet Kemal Kogak:
Insecta, taxonomy, nomenclature, fauna, ecology, catalogues, checklists of the Old World.

Expert referees according to the subject areas: Dr Peter Huemer (Austria) and Dr. Ole Karsholt (Denmark): Gelechioidea, and
some other Microlepidoptera groups in Palaearctic (Lepidoptera). Mag. Peter Buchner (Austria): Depressariidae (Lepidoptera). Dr J. B.
Heppner (U.S.A.): Microlepidoptera of Nearctic and Neotropical. Dr G. Baldizzone (Italy): Coleophoridae (Lepidoptera). Dr V. Korneyev
(Ukraine): Tephritidae, Pyrgotidae, Ulidiidae (Diptera). Prof. Dr Y.G.Verves (Ukraine): Sarcophagidae (Diptera). Dr Daniel Burckhardt
(Switzerland): Psyllidae (Homoptera). Prof. Dr E. Heiss (Austria): Hemiptera. Dr R. Ehrmann (Germany): Mantodea. Prof. Dr Mustafa
Unal (Bolu, Turkey): Orthoptera. Prof. Dr Hiiseyin Ozdikmen (Turkey): Coleoptera. Prof. Dr Suat Kiyak (Turkey): Hemiptera.

Plant taxonomy, flora and vegetation : Assoc. Prof. Dr Murat Unal, Asst. Prof. Dr Mesut Pmar, (Yuziincii Yil University, Van, Turkey).
Molecular studies : Asst. Prof. Dr Sibel Kizildag (Van Yiiziincii Yil University, Van, Turkey).

Editorial policy: the submitted manuscript is evaluated by the Chief Editor and Referee. In case of need, the manuscript is sent to
expert referees according to the subject areas.

| ALL RIGHTS RESERVED |

Correspondences should be addressed to: Prof, em Dr Ahmet Omer Kogak, c/o Yiiziincii Yil University, Fen Fakiiltesi,
Biyoloji Boliimii, Kampus, Van / Turkey. - e-mail: cesa tr@yahoo.com.tr

All the serials of the Cesa are archived regularly by the Internet Archive (300 Funston Ave., San Francisco, CA 94118,
U.S.A.), in accordance with the rules of the International Codes of Zoological Nomenclature (ICZN)

https://archive.org/

also recorded regularly by the Zoological Record, Thomson Reuters, Enterprise House, Innovation Way, Heslington,

York, YO10 5NQ, United Kingdom, https://clarivate.com/

8 http://www.cesa-tr.org/Pri.htm

9 http://www.cesa-tr.org/Miscell.htm

10 http://www.cesa-tr.org/Memoirs.htm

11 http://www.cesa-tr.org/CDF.htm

12 http://www.cesa-tr.org/Icon.htm

x 3 http://www.cesa-tr.org/CPAL.htm
^ http^/www.cesa-tr.org/Cesanews.htm

http://www.cesa-tr.org/Cesabooks.htm

27

Heterocera collection from northern Palawan ...